(Publications in scientific journals)

62. V. Cruz, Rey Benayas J.M., Ruiz-Beníto P., Villar-Salvador P. Restoration strategy modulates the effect of different shrub types on tree recruitment in Mediterranean forests (in preparation).

61. E. Andivia., Villar-Salvador P., Oliet, J., Puértolas J., Dumroese R.K, Ivetic V. Molina-Venegas R., Arellano E., Li G., Ovalle J. 2019 Size matters in forest restoration: a global meta-analysis on the relationship between the size of planted seedling and survival (in preparation).

60. L. Martínez-Baroja, Rey-Benayas J.M., Pérez Camacho L., Villar-Salvador P., 2019 Drivers of Quercus ilex establishment after 25 years of planted woodland islets to restore Mediterranean  old-fields (in preparation).

59. M. Uscola, Villar-Salvador P, Oliet J., Warren Ch. 2019 Growth rate across Mediterranean forest plants is linked to soil ammonium uptake preference. An overlooked trait in the fast growth syndrome? (in preparation)

58. M. Uscola, Villar-Salvador P., Jacobs D.F. 2019 Tolerance and recovery capacity after heat stress in pine species (in preparation)

57. P. Villar-Salvador P., Sánchez-Gómez D., Pérez-Camacho L., Cuevas J.A., García-Salgado G.J., Rey-Benayas JM. 2019. Oak species and structure of forest and shrubland drive forest recovery in old-fields: not all oaks get that far and so fast (in preparation).

56. A.O. Toca, Villar-Salvador P., Oliet J., Jacobs D.F. 2019 Root hydraulic standardization methods in seedlings determine the interpretation of nitrogen effects on pine species seedlings. (Submitted)

55. L. Martínez-Baroja, Pérez Camacho L., Villar-Salvador P., Rebollo S, Quiles P., Gómez-Sánchez D., Molina-Morales M, Leverkus A, Castro J, Rey-Benayas JM. 2018. Massive and effective acorn dispersal into open agroforestry systems by an overlooked vector, the Eurasian magpie (Pica pica). Ecosphere (in press).

54. V. Cruz, Villar-Salvador P., Ruiz-Benito P., Ibáñez I., Rey-Benayas J.M. 2019 Temporal dynamic of shrub facilitation shapes the mixing of evergreen and deciduous oaks in Mediterranean forest recovery. Journal of Ecology (in press).

53. A.O. Toca, Oliet J., Villar-Salvador P., Martínez R., Jacobs D.F. 2019 Ecologically distinct pine species show differential root development after outplanting in response to nursery nutrient cultivation. Forest Ecology and Management 451 (in press).

52. L. Fernández-Pérez, Zavala M.A., Villar-Salvador P., Madrigal-González J. 2019. Divergent last century tree growth along an altitudinal gradient in a Pinus sylvestris dry-edge population. Forests 10: 532 10.3390/f10070532

51. W. Shi, Villar-Salvador P., Li G., Jiang, X. 2019 Acorn size is more important than nursery fertilization for outplanting performance of Quercus variabilis container seedlings. Annals of Forest Science 72. https://doi.org/10.1007/s13595-018-0785-8.

50. V. Cruz-Alonso, Ruiz-Benito P., Villar-Salvador P., Rey-Benayas, J. M. 2019. Long-term recovery of Mediterranean forests depends on restoration strategy and forest type. Journal of Applied Ecology 56: 745-757. https://doi.org/10.1111/1365-2664.13340.

49. E. Andivia, Zuccarini P., Grau B, De Herralde F., Villar-Salvador P., Savé R. 2019. Rooting big and deep rapidly: the ecological roots of pine species distribution in southern Europe. Trees. Structure and Function 33: 293303 https://doi.org/10.1007/s00468-018-1777-x.

48. J. Wang, Villar-Salvador P., Li G. Y. Liu 2019. Moderate water stress does not inhibit nitrogen remobilization allowing high growth in high nitrogen content Quercus variabilis seedlings under drought conditions. Tree Physiology (in press; https://doi.org/10.1093/treephys/tpy130 ).

47. E. Andivia, Villar-Salvador P., Oliet J., Puértolas J., Dumroese K. 2019 How can my paper be useful for future meta-analysis on forest restoration plantations? New Forests 50: 255266.

46. E. Andivia, Madrigal-González J., Villar-Salvador P., Zavala, M.A. 2018. Does facilitation from adult conspecifics increase sapling resilience to repeated droughts in water-limited pine forest? Ecosphere 9, article e02282.

45. D. Salazar-Tortosa, Castro J., Villar-Salvador P., Viñegla B., Matías L., Michelsen A., Rubio de Casas R., Querejeta JI. 2018. The “isohydric trap”: a proposed feedback between water shortage, stomatal regulation and nutrient acquisition drives differential growth and survival of European pines under climatic dryness. Global Change Biology 24: 4069-4083

44. D. Salazar-Tortosa, Castro J., De Casas R.R., Viñegla-Pérez B., Sánchez-Cañete E.P., Villar-Salvador P. 2018. Gas exchange at whole plant level shows that a less conservative water use is linked to a higher performance in three ecologically distinct pine species. Environmental Research Letters 13: 045004.

43. L. Fernández-Pérez, Villar-Salvador P., Martínez-Vilalta J., Toca A.O., Zavala M.A 2018 Distribution of pines in Iberia Peninsula agrees with seedling differences in foliage frost tolerance, not with xylem embolism vulnerability. Tree Physiology 38:507-516.

42. W. Shi, Villar-Salvador P., Jacobs D.F., Li G., Jang X. 2018. Simulated predation of Quercus variabilis acorns impairs nutrient remobilization and seedling performance irrespective of soil fertility. Plant and Soil 423:295–306

41. A.O. Toca, Oliet J.A., Villar-Salvador P., Maroto J., Jacobs, D.F. 2018. Species ecology determines the role of nitrogen nutrition on the frost tolerance of pine seedlings. Tree Physiology 38: 96-108.

40. J. Castro, Molina-Morales M.,Leverkus A. B., Martínez L , Pérez-Camacho L, Villar-Salvador P, Rebollo S, Rey-Benayas J.M. 2017. Effective nut dispersal by magpies (Pica pica L.) in a Mediterranean agroecosystem. Oecologia 184: 183-192.

39. E. Andivia, Villar-Salvador P., Tovar L., Rabasa S., Rey-Benayas J.M. 2017. Multiscale assessment of woody species recruitrment in Mediterranean shrublands: facilitation and beyond. Journal of Vegetation Science 28: 639-648.

38. M. Uscola, Villar-Salvador P., Oliet, J. 2017 Root uptake of inorganic and organic N chemical forms in two coexisting Mediterranean forest trees. Plant and Soil 415:387-392

37. L. Matías, Castro J., Villar-Salvador P., Quero J.L., Jump A.S. 2017. Differential impact of hotter drought on seedling performance of five ecologically distinct pine species. Plant Ecology 218:201-212.

36. P. Villar-Salvador 2016. Restoration of Spanish pine plantations: A main challenge for the 21st century. Reforesta 1:53-66.

35. P. Villar-Salvador, Uscola, M. Jacobs, D.F. 2015. The role of stored carbohydrates and nitrogen in the growth and stress tolerance of planted forest trees. New Forests 46:813-839

34. José M. Rey Benayas, Martínez, L., Pérez-Camacho, L., Villar-Salvador, P., Holl, K. 2015. Predation and aridity slow down the spread of 21 year old planted woodland islets in restored Mediterranean farmland. New Forests 46:841-853.

33. M. Uscola, Villar-Salvador P., Gross, P., Maillard P. 2015 Fast growth involves high dependence on stored resources in seedlings of Mediterranean evergreen trees. Annals of Botany 115:1001-1013.

32. N. Heredia, Oliet J., Villar-Salvador P, Benito, Luis F, Peñuelas J.L. 2014. Fertilization regime interacts with fall temperature in the nursery to determine the frost and drought tolerance of the Mediterranean oak Quercus ilex L. Forest Ecology and Management 311:50-59.

31.M. Uscola, Villar-Salvador P, Oliet J., Warren C. 2014. Foliar absorption and root translocation of nitrogen from different chemical forms in seedlings of two Mediterranean trees. Environmental and Experimental Botany 104: 34-43.

30. M. Uscola, Oliet J., Villar-Salvador P., Díaz-Pinés E., Jacobs D. 2014 Nitrogen form and concentration interact to affect the performance of two ecologically distinct M editerranean forest trees. European Journal of forest Research 133:235–246.

29. P. Villar-Salvador, Peñuelas, J.L., Nicolás-Peragón, Fernando-Benito, L.F. Peñuelas, J.L. 2013 Is nitrogen fertilization in the nursery a suitable tool for enhancing the performance of Mediterranean oak plantations? New Forests 44:733-751.

28. P. Villar-Salvador; Peñuelas J.L., Jacobs D. 2013. Nitrogen fertilisation and drought hardening exert opposite effects on the stress tolerance of Pinus pinea L. seedlings Tree Physiology 33: 221-232.

27. P. Villar-Salvador, Puértolas J., Cuesta B., Peñuelas J.L; Uscola M., Heredia N., Rey Benayas, J. 2012. Increase in size and nitrogen concentration enhances seedling survival in Mediterranean plantations. Insights from an ecophysiological conceptual model of plant survival. New Forests. 43: 755-770.

26. D. Montesinos, Villar-Salvador P, García-Fayos P., Verdú, M. 2012. Genders in Juniperus thurifera have different functional responses to variations in nutrient availability. New Phytologist 193: 705-712.

25. B. Cuesta; Rey Benayas J.M., Gallardo A., Villar-Salvador P, González-Espinosa M. 2012. Soil chemical properties in abandoned Mediterranean cropland after succession and oak reforestation. Acta Oecologica 38: 58-65.

24. J. Climent, Pardos, M. Chambel, M.R., Lario, F., Villar-Salvador P. 2011. Biomass allocation and foliage heteroblasty in hard pine species respond differentially to reduction in rooting volume. European Journal of Forest Research 130: 841-850.

23. B. Cuesta, Vega J., Villar-Salvador P. Rey-Benayas JM. 2010. Root growth dynamics of Aleppo pine (Pinus halepensis Mill.) seedlings in relation to shoot elongation, plant size and tissue nitrogen concentration. Trees. Structure and Function 24:899908.

22. B. Cuesta, Villar-Salvador P, Puértolas J., Jacobs, D. Rey Benayas J.M. 2010 . Why do large, nitrogen rich seedlings better resist stressful transplanting conditions? A physiological analysis in two functionally contrasting Mediterranean forest species. Forest Ecology and Management 260:71-78.

21. J.M. Rey Benayas, Escudero A., Martín Duque J.F., Nicolau J.M., Villar-Salvador P, García de Jalón D., Balaguer L. 2010. A multiinstitutional Spanish master´s in ecosystem restoration: vision and four-year experience. Ecological Restoration 28:188-192.

20. J. G. Hodgson , M. Sharafi, A. Jalili, S. Díaz, G. Montserrat-Marti, C. Palmer, B. Cerabolini, S. Pierce, B. Hamzehee, Y. Asri, Z. Jamzad, P. Wilson, J. A. Raven, S. R. Band, S. Basconcelo, A. Bogard, G. Carter, M. Charles, P. Castro-Díez, J.H.C. Cornelissen, G. Funes, G. Jones, M. Khoshnevis, N. Pérez-Harguindeguy4, M. C. Pérez-Rontomé, F. A. Shirvany, F. Vendramini, S. Yazdani, R. Abbas-Azimi, S. Boustani, M. Dehghan, J. Guerrero-Campo, A. Hynd, E. Kowsary, F. Kazemi-Saeed, B. Siavash, P. Villar-Salvador, R. Craigie, A. Naqinezhad, A. Romo-Díez, L. de Torres Espuny and E. Simmons. 2010. Stomatal vs. genome size in angiosperms: the somatic tail wagging the genomic dog? Annals of Botany 105: 573584.

19. B. Cuesta, Villar-Salvador P, Puértolas J., Rey Benayas J.M. Michalet R . 2010 Facilitation of oak in Mediterranean shrubland is explained by both direct and indirect interactions mediated by herbs Journal of Ecology 98: 688-697

18. P. Villar-Salvador, Heredia N., Millard P. 2010 Remobilization of acorn nitrogen for early seedling growth in the Mediterranean oak Quercus ilex L., grown with contrasting nutrient availability Tree Physiology 30: 257-263

17. P. Villar-Salvador, Valladares F., Domínguez-Lerena, S., Ruiz-Diez, B., Fernández-Pascual, M., Delgado, A. and Peñuelas, J.L. 2008. Functional traits related to seedling transplanting performance in the Mediterranean leguminous shrub Retama sphaerocarpa: insights from a provenance, fertilization, and rhizobial inoculation study. Environmental and Experimental Botany 64:145-154.

16. J.M. Rey-Benayas, Cuesta B., Villar-Salvador P. and Jáuregui P. 2008 . Life-history traits in an evergreen Mediterranean oak respond differentially to previous experimental environments. Web Ecology 8: 74-83

15. S. Mollá, Villar-Salvador P . García-Fayos, P. Peñuelas, J.L. 2006. Physiological and transplanting performance of Quercus ilex L. (holm oak) seedlings grown in nurseries with different winter conditions. Forest Ecology and Management 237:218-226

14. P. Villar-Salvador, Puértolas, J., Planelles, R., Peñuelas Rubira, J. 2005. Effect of nitrogen fertilization in the nursery on the drought and frost resistance of Mediterranean forest species. Investigaciones Agrarias: Sistemas y Recursos Forestales 14:408-418

13. JG Hodgson, Montserrat-Marti G, Cerabolini B, Ceriani RM, Maestro-Martinez M, Peco B, Wilson PJ, Thompson K, Grime JP, Band SR, Bogard A, Castro-Diez P, Charles M, Jones G, Perez-Rontome MC, Caccianiga M, Alard D, Bakker JP, Cornelissen JHC, Dutoit T, Grootjans AP, Guerrero-Campo J, Gupta PL, Hynd A, Kahmen S, Poschlod P, Romo-Diez A, Rorison IH, Rosen E, Schreiber KF, Tallowin J, Espuny LD, Villar-Salvador P. 2005 . A functional method for classifying European grasslands for use in joint ecological and economic studies. Basic and Applied Ecology 6: 119-131.

12. JG Hodgson, Montserrat-Marti G, Tallowin J, Thompson K, Diaz S, Cabido M, Grime JP, Wilson PJ, Band SR, Bogard A, Cabido R, Caceres D, Castro-Diez P, Ferrer C, Maestro-Martinez M, Perez-Rontome MC, Charles M, Cornelissen JHC, Dabbert S, Perez-Harguindeguy N, Krimly T, Sijtsma FJ, Strijker D, Vendramini F, Guerrero-Campo J, Hynd A, Jones G, Romo-Diez A, Espuny LD, Villar-Salvador P. , Zak MR. 2005. How much wil it cost to save grassland diversity? Biological Conservation 122: 263-273.

11. P. Villar-Salvador, Planelles R., Enríquez E., Peñuelas Rubira J. 2004. Nursery cultivation regimes, plant functional attributes, and field performance relationships in the Mediterranean oak Quercus ilex L. Forest Ecology and Management 196: 257-266 .

10. P. Villar-Salvador , Planelles R., Oliet J., Peñuelas-Rubira J.L., Jacobs D.F., González M.. 2004 Drought tolerance and transplanting performance of holm oak ( Quercus ilex L.) seedlings after drought hardening in the nursery. Tree Physiology 24:1147-1155.

9. M. Verdú, Villar-Salvador P., García-Fayos P. 2004. Gender effects on the post-facilitation performance of two dioecious Juniperus species. Functional Ecology 18: 87-93.

8. S. Díaz, Hodgson, J.G., Thompson, K., Cabido, M., Cornelissen, J.H.C., Jalili, A., Montserrat-Martí, G., Grime, J.P., Zarrinkamar, F., Asri, Y., Band, S.R., Basconcelo, S. 1, Castro-Díez, P., Funes, G., Hamzehee, B., Khoshnevi, M., Pérez-Harguindeguy, N., Pérez-Rontomé, M.C., Shirvany, F.A., Vendramini, F., Yazdani, S., Abbas-Azimi, R., Bogaard, A., Boustani, S., Charles, M., Dehghan, M., de Torres-Espuny, L., Falczuk,,V., Guerrero-Campo, J., Hynd, A., Jones, G., Kowsary, E., Kazemi-Saeed, F., Maestro-Martínez, M., Romo-Díez, A., Shaw, S., Siavash, B., Villar-Salvador, P. & Zak, M.R. 2004 . The plant traits that drive ecosystems: Evidence from three continents. Journal of Vegetation Science 15: 295-304

7. J.H.C. Cornelissen, Cerabolini B., Castro-Díez P., Villar-Salvador P., Montserrat-Martí G., Puyravaud J.P., Maestro M., Werger M.J.A., Aerts R. 2003. Functional traits of woody plants: correspondence of species rankings between field adults and laboratory-grown seedlings? Journal of Vegetation Science 14: 311-322

6. F. Valladares, Villar-Salvador P ., Domínguez S., Fernández Pascual M., Peñuelas J.L., Pugnaire F.I. 2002. Enhancing the early performance of the leguminous shrub Retama sphaerocarpa (L.) Boiss.: fertilisation versus Rhizobium inoculation. Plant and Soil 240: 253-262

5. P. Villar Salvador, Ocaña L., Peñuelas J.L., Carrasco I. 1999. Effect of water stress conditioning on the water relations, root growth capacity, and the nitrogen and non-structural carbohydrate concentration of Pinus halepensis Mill. (Aleppo pine) seedlings. Annals of Forest Science 56: 459- 465.

4. P. Castro-Díez, Villar-Salvador P., Pérez-Rontomé C., Montserrat-Martí G. 1998. Leaf morphology, leaf chemical composition and stem xylem characteristics in two Pistacia (Anacardiaceae) species along a climatic gradient. Flora 193: 195-202.

3. P. Castro-Díez, Puyravaud J.P., Cornelissen J.H.C., Villar-Salvador P . 1998. Stem anatomy and relative growth rate in seedlings of a wide range of woody plant species. Oecologia 116: 57-66.

2. P. Villar-Salvador, Castro-Díez P., Pérez-Rontomé C., Montserrat-Martí G. 1997. Stem xylem features in three Quercus (Fagaceae) species along a climatic gradient in NE Spain. Trees-Structure and Function 12: 90-96.

1. P. Castro Díez, Villar-Salvador P ., Pérez Rontomé C., Maestro Martínez M., Montserrat Martí G. 1997 Leaf morphology and leaf chemical composition in three Quercus (Fagaceae) species along a precipitation gradient in NE Spain. 1997 Trees-Structure and Function 11: 127-134